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Abstract

Balanomorph cirripedes from the Eocene–Oligocene of the Hampshire Basin (United Kingdom) and the Middle Eocene of the Cotentin Peninsula, Manche (France) are described. A new genus, Vectibalanus, is founded, with the type species Balanus unguiformis J. de C. Sowerby, 1846; assigned to this are also Balanus erisma J. de C. Sowerby, 1846 and Vectibalanus mortoni sp. nov. In addition, a new species of Lophobalanus Zullo, 1984, L. fresvillensis sp. nov., is described. This is the first record of that genus from outside the eastern USA and the oldest species known to date. Cladistic analysis of 24 morphological characters suggests that Vectibalanus unguiformis is sister taxon to a group comprising the most derived balanomorph taxa, and thus represents an important transition in the evolution of the group, with the initiation of development of a complex parietal wall structure. Vectibalanus unguiformis was evidently adapted to low salinity habitats (10–30 ppt), and is the oldest known brackish water barnacle. The other species ( V. erisma, V. mortoni sp. nov.) occupied more clearly marine environments (>30 ppt). Balanomorph barnacles appeared simultaneously in the Priabonian (Upper Eocene) of the Gulf and Atlantic seaboards of the USA and northwest Europe, which probably represents a northerly migration from Tethys.
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Authors and Affiliations

Andrew Scott Gale
1 2

  1. School of the Environment, Geography and Geological Sciences, University of Portsmouth, Burnaby Building, Burnaby Road, Portsmouth PO13QL, UK
  2. Department of Earth Sciences, Natural History Museum, Cromwell Road, London SW75BD, UK
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Abstract

The uppermost Albian and lowermost Cenomanian succession at Abouda Plage, north of Agadir, in the Agadir Basin, western Morocco, is described in detail, and ammonites, microcrinoids and planktonic foraminifera are recorded and illustrated. The lower part of the Aït Lamine Formation yields ammonites indicative of the Pervinquieria (Subschloenbachia) rostrata and P. (S.) perinflata ammonite zones, and the Thalmanninella appenninica planktonic foraminiferan Zone. The base of the Cenomanian is identified at 42.2 m above the base of the Aït Lamine Formation, based on the lowest occurrence of the planktonic foraminiferan Thalmanninella globotruncanoides Sigal, 1948. Lower Cenomanian ammonites of the Graysonites adkinsi Zone enter 3 m higher in the succession. Microcrinoid zones AlR11 and AlR12 are identified in the Upper Albian, and the base of the CeR1 Zone coincides with the lowest occurrence of Cenomanian ammonites. The ammonite and microcrinoid occurrences and detailed distributions are very similar to those found in north central Texas, which, in the Cenomanian, was 5,300 km to the west. The new records suggest that the G. adkinsi Zone is equivalent to the uppermost (Lower Cenomanian) part of the Pleurohoplites briacensis Zone of the Global Stratotype Section for the base of the Cenomanian stage. An hiatus, of global extent, immediately underlies the base of the G. adkinsi Zone and is represented in the Agadir Basin by an erosion surface containing bored and encrusted hiatus concretions.
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Authors and Affiliations

Andrew Scott Gale
1 2
William James Kennedy
3 4
Maria Rose Petrizzo
5

  1. School of the Environment, Geography and Geological Sciences, University of Portsmouth, Burnaby Building, Burnaby Road, Portsmouth PO13QL UK
  2. Earth Science Department, Natural History Museum, Cromwell Road, London SW75BD, UK
  3. Oxford University Museum of Natural History, Parks Road, Oxford, OX13PW
  4. Department of Earth Sciences, South Parks Road, OX13AN UK
  5. Dipartimento di Scienze della Terra ‘A. Desio’, Università degli Studi di Milano, via Mangiagalli 34, 1-20133 Milano, Italy
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Abstract

The stratigraphy of the upper Fredericksburg and lower Washita groups of northern Texas and southern Oklahoma is described, and biostratigraphical correlation within the region, and further afield, using micro­ crinoids, ammonites, planktonic foraminiferans and inoceramid bivalves is summarised. The taxonomy of the roveacrind microcrinoids is revised by the senior author, and a new genus, Peckicrinus, is described, with the type species Poecilocrinus porcatus (Peck, 1943). New species include Roveacrinus proteus sp. nov., R. morganae sp. nov., Plotocrinus reidi sp. nov., Pl. molineuxae sp. nov., Pl. rashallae sp. nov. and Styracocrinus thomasae sp. nov. New formae of the genus Poecilocrinus Peck, 1943 are Po. dispandus forma floriformis nov. and Po. dispandus forma discus nov. New formae of the genus Euglyphocrinus Gale, 2019 are E. pyramidalis (Peck, 1943) forma pyramidalis nov., E. pyramidalis forma radix nov. and E. pyramidalis forma pentaspinus nov. The genera Plotocrinus Peck, 1943, Poecilocrinus and Roveacrinus Douglas, 1908 form a branching phylogenetic lineage extending from the middle Albian into the lower Cenomanian, showing rapid speciation, upon which a new roveacrinid zonation for the middle and upper Albian (zones AlR1–12) is largely based. Outside Texas and Oklahoma, zone AlR1 is recorded from the lower middle Albian of Aube (southeastern France) and zones AlR11–CeR2 from the Agadir Basin in Morocco and central Tunisia. It is likely that the zonation will be widely applicable to the middle and upper Albian and lower Cenomanian successions of many other regions.
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Authors and Affiliations

Andrew Scott Gale
1 2
Jenny Marie Rashall
3
William James Kennedy
4 5
Frank Koch Holterhoff
6

  1. School of the Environment, Geography and Geological Sciences, University of Portsmouth, Burnaby Building, Burnaby Road, Portsmouth PO13QL UK
  2. Earth Science Department, Natural History Museum, Cromwell Road, London SW75BD, UK
  3. Department of Earth and Environmental Sciences, University of Texas at Arlington, 76019 USA
  4. Oxford University Museum of Natural History, Parks Road, Oxford, OX13PW
  5. Department of Earth Sciences, South Parks Road, OX13AN UK
  6. 1233 Settlers Way, Lewisville, TX 75067 USA

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