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Abstract

The first recognition of a tracemaker responding to a temporary shift in the redox boundary is recognized. This is recorded by a new trace fossil, Sursumichnus orbicularis igen. et isp. nov., which is established for mound-like structures on the upper surfaces of sandstone beds from the Borjomi Flysch (upper Paleocene–lower Eocene) in the Lesser Caucasus (Georgia). It is connected with the spatangoid echinoid burrow Scolicia de Quatrefages, 1849 and interpreted as a resting trace of the same tracemaker produced after moving up from a deeper position within the sediment. The resting is caused by an episode of unfavourable conditions related to shallowing of the redox boundary. The trace fossil is a component of the Nereites ichnofacies.
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Authors and Affiliations

Alfred Uchman
1
Zurab Lebanidze
2
Nino Kobakhidze
3
Tamar Beridze
3
Davit Makadze
4
Koba Lobzhanidze
3
Sophio Khutsishvili
3
Rusudan Chagelishvili
5
Kakha Koiava
2
Nino Khundadze
4

  1. Faculty of Geography and Geology, Institute of Geological Sciences, Jagiellonian University, Gronostajowa3a, 30-387 Kraków, Poland
  2. Department of Geology, Faculty of Exact and Natural Sciences, Iv. Javakhishvili Tbilisi State University, University str. 13, 0186 Tbilisi, Georgia
  3. Alexander Janelidze Institute of Geology, Iv. Javakhishvili Tbilisi State University, Politkovskaia 31, 0186 Tbilisi, Georgia
  4. Alexander Tvalchrelidze Caucasian Institute of Mineral Resources, Tbilisi State University, 12 Mindeli str., 0186 Tbilisi, Georgia
  5. Department of Geology and Paleontology, Georgian National Museum, 3 Purtseladze str., 0105 Tbilisi, Georgia

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